Ouattara, Salimata, West African Scientific Service Centre on Climate Change and Adapted Land Use (WASCAL), Graduate Research Program on Climate Change and Disaster Risk Management, Kwame Nkrumah University of Lomé, Lomé, Togo Laboratoire des Milieux Naturels et Conservation de la Biodiversité, U.F.R. Biosciences, Université Félix Houphouët-Boigny, Côte d’Ivoire, Côte d'Ivoire